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The Giant Turtle was seen off the coast of Alaska and sunk the ship that Nick and Audrey were having a romantic cruise on. It attacked again when it found them on a nearby deserted island, but they were saved by Komodithrax. When it attacked both Komodithrax and Zilla's nest, it fell into a crater with Komodithrax.
It is a reference to both Gamera (he walked on all fours in his first two films) and Kamoebas (Kamoebas had a neck that could stretch out) (Beacause of the spiky shell some fans speculate it could be a reference to Anguirus as well).