MUTO; Massive Unidentified Terrestrial Organism. „ 

William Stenz (Godzilla)


The MUTOs (ムートー,   Mūtō?) are giant parasitic daikaiju created by Legendary Pictures that first appeared in the 2014 film, Godzilla, as the primary antagonists. A third MUTO appears in the sequel Godzilla: King of the Monsters, as a minor Titan that obeys Ghidorah and later Godzilla.


The MUTO's name is an acronym for "Massive Unidentified Terrestrial Organism."[6] According to Godzilla: Awakening, this is a designation Monarch gives to all giant monsters as a sort of placeholder name. In this comic, both Godzilla and Shinomura were known as MUTOs before being given their respective names.

While initially lacking an official Titanus designation, the opening of Godzilla vs. Kong designates them as "Titanus MUTO", which is inaccurate given the opening had many inconsistencies due to production delays.[7] Officially, they are classified as Titanus Jinshin-Mushi and are the same species as their progenitor, MUTO Prime, according to Arvid Nelson.[1]

In one of the original screenplays for Godzilla, the MUTOs were called "Hokmuto" and "Femuto," which were short for "Hokkaido MUTO" and "Female MUTO," respectively.[8]

In the final film, the MUTOs are referred to as such but are also referred to numerically, with the male MUTO being designated as "MUTO 1" and the female as "MUTO 2" respectively on a monitor present near William Stenz. The third MUTO that was seen in Godzilla: King of the Monsters has been given the official title of "Queen MUTO", and also given the nickname "Barb" by the director and writer of Godzilla: King of the Monsters, Michael Dougherty.[9]



The MUTOs are known to have been developed from Rokmutul and Pterodactyl, two early monsters from the development of Godzilla. In the Comic-Con 2012 teaser trailer for the film, an unnamed, tardigrade-like, multi-legged monster was created to confirm that Godzilla would be fighting another monster in Legendary Pictures' Godzilla.

According to witnesses of the Godzilla trailer shown in Comic-Con 2013, the MUTOs were originally spider-like creatures with long, thin limbs and scythes. This changed in the actual film.

One of the original screenplays of Godzilla had the male MUTO emerge from its chrysalis in Hokkaido, Japan, instead of the fictional city of Janjira. The female MUTO still appeared in Nevada, however.[8]

The MUTOs went through several dozen concepts before Legendary settled on their final designs. Some concepts had the male MUTO with four wings,[10] and others had him with only six total appendages (including wings).[11] Many of the early designs featured a rougher-looking, more 'organic' appearance, rather than the smooth and metallic-looking final design.


The MUTO's bodies are covered in an iridescent, metallic grayish-black exoskeleton. Notably, they have narrow red, visor-like eyes. The shape of their head is reminiscent of Gyaos, and they have triangular jaws resembling a hooked beak.

The male MUTO's body, showing a slim, airborne Titan.

As a species, the MUTOs are sexually dimorphic, as there are several notable differences between males and females. The female MUTO has two main pairs of forelimbs and a smaller pair on her abdomen, and is much bigger than the male MUTO, standing nearly twice the height of the male and being almost as tall as Godzilla. She also has a pouch-like structure on her lower abdomen, which while gravid visibly contains the glowing eggs. The male MUTO is similar in appearance, except that he is much smaller with a more nimble build, and his front forelimb pair has been naturally repurposed to form his wings, which are long, pointed and membranous, similar to those of a pterosaur. Both males and females possess long and slender hind limbs that are digitigrade in structure, with flat, broad feet ending in two hoof-like toes. The forelimbs all end in two digits and take on a pseudo claw form, being curved like a sickle. When walking, the MUTOs use the front "knuckle" of their claws, giving it a hook-like appearance. Female MUTOs, under certain circumstances, can morph into a MUTO Prime, further adding to the differences between males and females of the species.

The female MUTO's torso, head, limbs, and eggs.

MUTO 3, referred to as the Queen MUTO, looks slightly different in appearance to the female MUTO from 2014. This new MUTO has numerous spike-like protrusions running down her back that indicate older age, as well as leg joints that all end in a noticeable point. Unlike her deceased kin, the Queen MUTO's exoskeleton is more of a gray color instead of black. Her body is covered in scars from past battles and mating rituals.

MUTO eggs are spherical in shape and have a bright orange bioluminescent hue. They are almost completely transparent, revealing the developing MUTO embryo inside, which is spider-like in appearance. MUTO eggs appear to be encased in a ootheca-like structure when laid. MUTO cocoons are made from a hardened, rock-like substance and notably curve inward the taller it gets. Segments of the cocoon produce bright orange bioluminescent flashes, occasionally revealing the silhouette of the maturing adult within.


The MUTOs are portrayed by CGI.


The roar first appeared as an MP3 file hidden in the official Godzilla site.[12] The roar can be heard when the official site loads up, though it is faint due to Godzilla's roar being much louder than the MUTOs' roar.

The MUTOs have very unique roars, many of which are deep and blaring with occasional crackles and snapping noises. At a few points, the MUTOs make trumpet-like cries and groans similar to creaking doors or Geiger counters. The male and female vocalizations are slightly different; the male emits higher-pitched calls and shrieks, while the female has deeper, more guttural roars. Both MUTOs emit distinctive chirps and clicks when they touch snouts as a courtship greeting.


As individuals, the MUTOs aren't given much character development. Although they exhibit intelligence like the other Titans, their sole goals revolve around survival, radiation consumption, and reproduction.

The two MUTOs that were awakened in 2014 are usually very indifferent to humans and manmade structures around them, and they almost completely shrugged off blasts and projectiles when the military attempted to engage them, similarly to their rival Godzilla. But compared to Godzilla, who is likely to navigate around larger manmade structures in his way and otherwise avoid causing significant damage if he can, the MUTOs are almost completely indifferent: they will go straight through large manmade obstacles in their way without any apparent notice or regard for the humans their actions harm or inconvenience. Some examples of this include the destruction the male MUTO caused whilst escaping at the Janjira nuclear plant site, the destruction the male carelessly causes in San Francisco before mating, and the female MUTO travelling straight through Las Vegas in a continued straight line (brushing through skyscrapers and other landmarks) instead of navigating around it. However, the male MUTO displayed some curiosity at Honolulu, when lights on a train track reactivating drew its attention, and it bit down on a train passing along the track. The male also appeared to deliberately utilize its EMP blast against a group of fighter jets approaching its position, unlike the female who ignored such attacks at Las Vegas.

The MUTOs display courtship

When the 2014 MUTOs met up to mate, the male provided the female a nuclear warhead it had promptly seized as a sign of courtship, and they appeared to display affection by pressing their heads together once the female accepted the courtship sign. Afterwards, the male MUTO seemingly acts as a sentry, guarding the San Francisco area they've claimed as their nest against Godzilla whilst the female reproduces, although the female immediately afterward didn't hesitate to charge into a fight with Godzilla herself when the latter got past the male to her nest. The male and female MUTOs display teamwork in their fight against Godzilla, particularly when they gain an upper-hand and begin beating him down. The male also appears to display protectiveness of the female based on its attacks against Godzilla, such as dragging Godzilla off of the female. The male takes advantage of its streamlined build and flight, favoring surprise strikes and evasion where possible when engaging Godzilla.

The male MUTO drags Godzilla away from the female. This allows the latter to pick herself off the ground.

When the MUTOs' nest was destroyed, the female immediately noticed and charged in a panic to the site, displaying strong maternal instinct; whilst the male seemed to only notice when the female reacted, but promptly abandoned its fight with Godzilla to follow her. The female MUTO immediately dug and salvaged through the explosion site for any of her eggs and notably mourned when she found they'd all perished. After her young's death, the female MUTO promptly went berserk with rage, abandoning the fight with Godzilla and charging after the halo team, notably going out of her way to slaughter them with her jaws and arms whilst seemingly focusing on the boat holding the nuclear warhead they'd stolen. Whether the female targeted the team because she knew they'd caused the explosion that destroyed her nest, or because she was directing her rage at the first thing she saw after her young's death (Ford Brody had notably caught her attention right after her nest's destruction) is unknown.

Despite the actions and hostility of the 2014 MUTO pair, the Queen MUTO that was awakened in 2019 apparently conforms to the Titans' hierarchy to some extent, and is consequently mutually tolerant of Godzilla upon bowing to him as the new Alpha after he defeats Ghidorah.


The MUTOs are ancient parasitic Titans that existed during the Permian period of Earth's history, and are primarily driven by reproduction. The artbook Godzilla: The Art of Destruction states that the MUTOs reproduced by killing members of Godzilla's species and laying eggs inside their prey's radioactive carcasses. In Godzilla: Aftershock, Dr. Emma Russell theorizes based on analysis of MUTOs' reproductive organs that if two MUTOs successfully reproduce and the brood matures, the matured spawn will destroy, dominate, and reshape entire ecosystems to their own needs. Once that has been accomplished, she theorizes that the MUTOs would then turn on each other, with only the strongest individual surviving. The graphic novel also introduces the MUTO Prime, a matriarch-like MUTO that specializes in luring out and overpowering members of the Titanus Gojira species before implanting MUTO eggs in their stomach linings via tendril-like ovipositors. Though the MUTO Prime needs the Titanus Gojira to remain alive as an incubator, after the eggs have been laid, they slowly feed on the Titan host's nuclear-rich hemoglobin which eventually kills the host, and the eggs continue to gestate for a matter of centuries. Like Godzilla, the MUTOs feed off of the Earth's natural radiation, but can adapt and feed off of man-made radiation, such as warheads and nuclear reactors.

In Aftershock, several male MUTOs could be seen terrorizing an ancient human civilization, showing that the species was active during the time of early humans. Geological dating on MUTO egg chambers corresponds with at least two mass extinctions in Earth's history, heavily implying that the MUTOs were responsible for said extinction events. At some point, the majority of the MUTOs died off, save for two eggs that were injected by a MUTO Prime into the body of Dagon, a member of Godzilla's species, in 1100 BCE. The two eggs fed off of Dagon's radiation-rich blood, which eventually killed him. Afterwards, the eggs remained alive, intact and dormant inside Dagon's body for millennia.



The male MUTO's spore

The MUTO eggs in Dagon's skeleton were discovered in the Philippines in 1999 after a mining operation unknowingly drilled into a cave containing the skeleton. By the time Monarch investigated, one of the eggs had already hatched a male MUTO, catalyzed by the mining disturbance (which in turn caused the radiation levels in the cave to spike), and the creature had escaped to sea at the nearby coastline, whilst the second egg containing a female remained dormant. After two weeks[13], the newly-hatched male MUTO burrowed his way to the Janjira nuclear power plant in Japan, where he caused the plant to collapse from underneath, killing Sandra Brody and several other workers, and causing the entire area to be evacuated because of radiation leaking from the plant. Despite the destruction of the plant, the MUTO attached himself to the reactor and entered a cocoon-like state, absorbing all the radiation from the surrounding area. The second pod containing the female MUTO was vivisected and studied for years by Monarch, and once Serizawa confirmed the female was inert, the egg was taken by the Americans to the Yucca Mountain Nuclear Waste Depository in Nevada.

The male M.U.T.O. at the Honolulu airport

For the next 15 years, Monarch maintained a research base in the ruins of the Janjira plant, where they observed and studied the MUTO while it fed on the reactor. As the reactor was depleted and the MUTO matured, it began giving off electromagnetic pulses which interfered with Monarch's equipment. Ishiro Serizawa, who was in charge of the operation, regretfully gave the order to kill the MUTO before it could destroy their base. The cocoon was destroyed with electrical currents released from cables surrounding it. A group of armed operatives approached the remains of the cocoon to confirm the MUTO was destroyed, but the creature was still alive and attacked them. The MUTO laid waste to the power plant once again before sprouting his wings and taking off. The United States Navy, under the command of Admiral William Stenz, began to pursue the MUTO as it crossed the Pacific Ocean. Eventually, the male MUTO arrived in Hawaii, where it had dragged a Russian nuclear sub ashore and was feeding on its torpedoes. A group of soldiers were sent to investigate, but were attacked by the MUTO. Fighter jets were sent in to try and stop the creature, but the MUTO released an EMP, causing the jets to fall out of the sky and crash. The MUTO proceeded to terrorize the Honolulu airport but was confronted by Godzilla, who had come ashore to hunt it. The two monsters briefly battled until the MUTO retreated and flew over the ocean, with Godzilla in pursuit.

The female MUTO briefly looks in on a bar she had torn open.

Meanwhile, Serizawa and his assistant Vivienne Graham concluded that the male MUTO was using echolocation to signal to the female MUTO that he was mature and ready to mate. Serizawa worried that the spore containing the female was not dormant, and the military sent a team to the waste depository to investigate. When the soldiers arrived, the facility was destroyed and the female MUTO had already broken out. The female MUTO terrorized the city of Las Vegas and began to head west to California. The military formed a plan to lure both MUTOs and Godzilla out to sea with a nuclear warhead, then detonate it in an attempt to kill all three. Two warheads were transported by train, but the female MUTO intercepted it in the California wilderness, killing all the personnel except for Ford Brody and eating one of the warheads. The other warhead was recovered and airlifted to San Francisco Bay, where it was armed, but it was stolen by the male MUTO, who presented it to the female in downtown San Francisco.

As a pair, the two MUTOs are able to easily overpower Godzilla.

After acquiring the warhead, the female MUTO created a nest and attached her eggs to the warhead, nourishing them with the radiation. The male meanwhile attempted to distract Godzilla, who had broken through the Golden Gate Bridge and come ashore. Eventually, Godzilla reached the nest and took on both MUTOs at once, while an extraction team led by Ford Brody arrived to recover the armed warhead. The bomb was removed from the nest, and Ford ignited a gas explosion to destroy the MUTO eggs. The explosion attracted the attention of the MUTOs, who had overpowered Godzilla, and they went to investigate. The female MUTO was stricken with grief after seeing her young killed but became enraged when she saw Ford, the man responsible. Before the MUTO could kill Ford, Godzilla emerged from behind her and pummeled her mercilessly with his atomic breath. The male MUTO attacked Godzilla from behind, allowing the female to pursue Ford and his team as they attempted to take the bomb out to sea. When the male MUTO attempted to strike Godzilla from behind once again, Godzilla anticipated it and smacked the M.U.T.O. with his tail, impaling him on a building and killing him.

The male MUTO is tricked into flying into Godzilla's trap.

The female MUTO is killed by Godzilla's "kiss of death"

Ford's team arrived at the docks with the bomb but were all slaughtered by the female MUTO, leaving Ford the only one left. Ford grabbed the bomb and placed it on a boat. Unable to defuse it, he started the boat in an attempt to take it out over the bay before it could detonate. However, the female MUTO's EMP field disabled the boat, leaving her face-to-face with Ford. Ford drew his pistol and aimed it at the MUTO, but knew he stood no chance and closed his eyes as she prepared to kill him. Suddenly, the female MUTO was pulled back by Godzilla, who pried open her jaws and fired his atomic breath down her throat, severing her head from her body and killing her.

Godzilla: Aftershock

In a flashback, both the male and female MUTOs can be seen as eggs after being injected by a MUTO Prime into the stomach lining of Dagon. The parasitic infection eventually kills the Gojira, while the two MUTO eggs continue to gestate.

Dr. Emma Russell discusses the MUTOs during a meeting with her colleagues at an outpost built over Dagon's skeleton, theorizing about their lifecycle and affect on the world. After studying the geological dates of MUTO egg chambers and Phoenician tablets, she discovers that the parasitic Titans were responsible for several of Earth's mass extinctions and even a "global dark age" that wiped out countless civilizations and cultures. This is confirmed by another flashback showing several male MUTOs terrorizing an unknown ancient civilization.

Using the bioacoustics of the MUTOs, known as the "Drums of Raijin" to the ancients, Emma creates an ORCA prototype in order to distract MUTO Prime long enough for Godzilla to defeat her.

Godzilla: King of the Monsters

The second female MUTO being pacified by the ORCA during its rampage in a news headline

Following the battle of San Francisco, the severed head of the female MUTO was recovered and stored at Castle Bravo, the outpost where Monarch studies Godzilla, along with the male MUTO and MUTO Prime's corpses.

A second female MUTO, nicknamed "Queen MUTO" due to her unique dorsal spikes resembling a crown, was among the Titans being awakened by Ghidorah's call from her slumber deep beneath a strip mall in Hoboken, New Jersey. Following her new alpha's orders, the MUTO begins to hunt on Earth with her new pack by attacking the city of Hoboken.

The Queen MUTO bowing to Godzilla in submission

However, she later hears Madison Russell activate the ORCA in Boston, causing her to become docile. The Queen MUTO makes the journey there but does not move quickly enough to arrive before Godzilla kills Ghidorah. Following Behemoth and Scylla, the MUTO confronts Godzilla. After Rodan bows to Godzilla, the Queen MUTO also bows, followed by Scylla, Behemoth, and Methuselah, acknowledging Godzilla as the new alpha of the Titans.

Godzilla Dominion

Though she doesn't appear in the graphic novel, Queen MUTO, alongside other Titans, are sent into dormancy by Godzilla.

Kingdom Kong

As fighter pilot Audrey Burns' squadmates attempt to comfort her over her guilt in not stopping Camazotz when he first awakened, several bring up losing friends and family to Titans as a way to relate to her. One mentions losing his mother when the MUTOs rampaged through San Francisco back in 2014.

Godzilla vs. Kong

Queen MUTO remains dormant during Godzilla's search for Ghidorah's remains that were desecrated by Apex Cybernetics and for his fight against Kong and later, Mechagodzilla, as per her alpha's instruction.


Bio-Atomic Nature

Monarch classifies the MUTOs as Bio-Atomic in nature[14], as they are able to feed off of nuclear energy, gain strength, and harness power from it.


Both male and female MUTOs are armed with hook-like claws that are used to hack at and stab any Titan aggressors. The MUTOs used their claws to stab Godzilla during the battle in San Francisco, with these attacks being strong enough to draw blood.


Both MUTOs were durable enough to take barrages of both small and large arms fire without any damage. However, the attacks from Godzilla are shown to harm them greatly, as both the male and female screeched in pain whenever they were bitten or clawed by him. The female was able to survive being blasted by Godzilla's atomic breath with no visible damage.


Both MUTOs can use echolocation to communicate, find sources of radiation, and locate each other.

Electromagnetic pulse

The male MUTO's EMP charge

The male MUTO can unleash electromagnetic pulses from his claws, which can disable electrical appliances in a five-mile-wide radius (though it is also implied he does not necessarily have to rely on his claws to unleash an EMP, as when he gets the warhead, he unleashes it from within mid-air). The female can instead surround herself with an EMP field (labeled the "Sphere of Influence" by the news) which performs the same function. There doesn't appear to be any limits to how often either MUTO can use their EMP abilities. In the official novelization, it is explained the EMP ability evolved as a defense mechanism, used to prevent Godzilla's species from using their atomic breath. In an earlier screenplay and also in the final film's novelization, it's explained the MUTOs' EMP weakens Godzilla's atomic breath and renders him unable to use it by dampening the bio-electric spark which ignites it - it's never explicitly confirmed if this is canon in the film, but Godzilla's atomic breath does appear weaker when fighting the MUTOs than when he later fights Ghidorah and other opponents.


Despite their destructive nature, the MUTOs passively produce life-giving radiation that exponentially increases plant growth and other forms of life, a trait that many Titans are known to have. Las Vegas is seen to have transformed into a lush jungle after the female MUTO had passed through it in 2014. San Francisco shares a similar fate after both MUTOs and Godzilla battled in the former city.


The male MUTO has wings that he can use to fly, but his top speed is undetermined. Naturally, these wings aided him in seeking out radiation to offer to the female. The wings also gave the male a massive combat advantage against Godzilla, being able to ambush him from the skies with a series of strafing and grappling attacks. He would usually be able to escape back into the skies before Godzilla could retaliate.


The male MUTO ambushes Godzilla, distracting him from finishing off his mate.

The MUTOs have shown to be intelligent enough to work as a team as the male MUTO was able to drag Godzilla away from the female MUTO, who then attacked him in unison. The female was also able to quickly determine Ford's involvement in destroying her nest. The male MUTO could be seen deliberately using his EMP attack to disable fighter jets that were attacking him.

Upon witnessing Godzilla’s victory over Ghidorah, the Queen MUTO was intelligent enough to not challenge the former and decided to yield to her new alpha.


A group of MUTO eggs latched around the warhead

The will to reproduce for MUTOs is their main driving force and is so rapid that it is described as a "breeding force" that can very quickly destroy the global ecosystem. As parasites, the MUTOs' standard forms are born of eggs implanted by a MUTO Prime (itself believed to be a later stage in the standard female's life cycle) inside a Titanus Gojira specimen. The female MUTO can reproduce sexually - and unlike the MUTO Prime, which implants a seemingly small handful of large, opaque and spore-like eggs inside a live Titanus Gojira host, the female which emerged in 2014 was instead observed laying hundreds of smaller, translucent eggs at once; doing so mere minutes to hours after her eggs were fertilized by contact with the male MUTO, creating an improvised nest by smashing a large crater into the earth and laying the eggs around a secondary source of radiation. While the carcass of a Titan is preferred, other radioactive sources can be used to gestate the eggs, as seen with the breeding pair from 2014 using a warhead to do so.


Female MUTOs have the ability to molt into a MUTO Prime, granting them increased size and strength, along with seismic concussive powers and other abilities. Dr. Emma Russell theorizes that this transformation takes place after a female MUTO kills and consumes her mate, or after killing off her broodmates. Arvid Nelson claims that if a female finds herself in isolation from other members of the species, the transformation can occur,[1][15] though at the same time the reasons are complex and variable.[16]

Speed and agility

The male MUTO evades Godzilla

The male MUTO was able to outmaneuver Godzilla due to his wings and smaller size, while the female MUTO possessed enough speed to outrun humans. The male's smaller size also allowed him to climb and perch atop buildings with ease.


The MUTOs were shown to have excellent stamina and resilience. The male spent most of his time after hatching flying around in a constant hunt for radiation, and most importantly, a potential mate. In his final confrontation in San Francisco, the male fought Godzilla for several hours by himself, from afternoon to well into the night, while the female built a nest for their eggs. The female also displays excellent stamina, traveling from Nevada to San Francisco in the span of a couple of days, tearing through any obstacles between her with her massive size. She fought alongside the male in San Francisco, even while being incredibly injured from Godzilla's attacks, most notably his atomic breath.

Strength and combat

Both the male and female MUTOs possess immense physical strength. The male MUTO was strong enough to drag Godzilla away from the female MUTO without difficulties and dredge up a Russian attack submarine from the depths of the ocean and pull it dozens of miles inland on the island of Hawaii; the fact that he could perform such a feat, and later attack a ship in San Francisco Bay to collect a nuclear warhead, implies that he possessed some swimming capabilities. The female was able to claw herself out of Mount Yucca, as well as being able to destroy buildings with ease. She was also able to toss and ram Godzilla around, despite his immense size and weight.


Inner mouth

While the MUTO's exoskeletons were durable enough to withstand a point-blank atomic breath blast, the insides of their mouths and throats are not armored as such, and a blast of Godzilla’s atomic breath into the female's mouth easily decapitated her.

Parental instinct

The MUTOs are excellent parents and were fiercely protective of their eggs. Though the two MUTOs had tag-teamed Godzilla, having gotten the upper hand and were about to kill him, the sight of an explosion near their nest immediately caused them to both abandon the fight, which gave Godzilla enough time to recover and charge up his atomic breath.

Relative fragility

Godzilla easily spins the male MUTO around before sending him flying.

The male MUTO, despite being able to evade all of Godzilla's attacks with his speed and agility, was killed after being struck by the latter's tail into a building and getting impaled by the rubble.

Video games

Godzilla: Smash3

MUTO appears in Godzilla: Smash3 as an enemy that Godzilla fights against.


  • The MUTOs' role in Legendary's Godzilla mirrors that of the Gryphon from the scrapped 1994 American Godzilla film.
    • In the unmade script, Godzilla is viewed as a threat by the humans but is eventually realized to be a lesser evil when a more dangerous monster (who is Godzilla's enemy) attacks. The script also has Godzilla decapitating the Gryphon with his breath, and the humans allow him to leave in peace, just like in the 2014 film.
    • They are also similar to the Queen Bitch from the unmade sequel to the 1998 film. Both are insect-like kaiju whose populations are kept in check by the predation of Godzilla's species.
  • The MUTOs are the first American-made kaiju to be specifically created for a Godzilla film, not counting the Gryphon from the unmade 1994 American Godzilla film and the TriStar Godzilla from the 1998 American film.
  • Strangely for Godzilla antagonists, the MUTOs are portrayed with a hint of sympathy for their plight: they are not outright evil or malicious creatures, merely animals trying to survive and raise a family in a strange hostile world, a fact emphasized by scenes of the affectionate courtship or the mother mourning the destruction of her young.
  • Despite their resemblances to arthropods, the MUTOs possess vertebrate biology.
  • The MUTO's behavior combines aspects from various real-life insects:
    • The male remains dormant for 15 years before emerging as a winged adult, similar to a periodical cicada.
    • The female is twice the size of the male but lacks the wings, like fireflies and certain wasps.
    • They use long-distance mating calls, similar to crickets.
    • They grow and mature inside living hosts like endoparasitoid wasps do.
    • The female lays her eggs on a readily-prepared food source, as flies and beetles do.
    • The courtship is somewhat like that of some spiders and mantises, where the male presents the female with a nuptial gift.
  • The production crew were readjusting the MUTO's design in the 2014 film until the very last minute.[17]
  • According to Godzilla: The Official Movie Novelization:
    • The MUTOs' EMP renders guidance systems unable to lock onto them from afar.
    • It's only implicit in the film, but Ford Brody inwardly speculates that the MUTOs' electromagnetic influence affecting the weather is causing the thunderstorm that rolls over San Francisco after they set up nest.
  • In Rogue One: A Star Wars Story, in Saw Gerrera's cave, paintings of the MUTOs can be spotted. Gareth Edwards directed both Godzilla and Rogue One, leading to the cameo.
  • Some aspects of the MUTO's life cycle are unclear:
    • The male was said to have emerged from the spore as a larva, yet the developing young seen inside the eggs already resembled the full-grown adults.
    • While the female remained dormant the whole time, the male emerged as a larva and pupated before emerging as an adult: in contrast, the female emerges from the spore already fully grown, seemingly having skipped the larva and pupa stages. It is likely that she hatched, and then pupated while in Yucca Mountain.
    • Despite not having met up with the male yet, the female MUTO emerged from Yucca Mountain already pregnant.
    • In Godzilla: Aftershock, Dr. Emma Russell theorized that the female MUTO would have eaten the male after mating and morphed into a MUTO Prime, yet this did not happen, as the two MUTOs could be seen practicing cooperative breeding instead.
  • Though the female MUTO simply crouches into the nest when she lays her eggs, the eggs are neatly arranged in rows when Ford and his team enter the nest. She may possess an ovipositor that allows her to adjust her eggs to the source of energy.
    • This is further implied by the fact that MUTO Prime possesses several.
  • The MUTOs are some of the few kaiju to almost kill Godzilla.
  • Michael Dougherty added a MUTO to Godzilla: King of the Monsters because he liked the design of the species and loved the idea that there are still other living MUTOs in the world.[18]
  • The MUTOs resemble the monster from the 2008 film Cloverfield.

In other languages

  • Russian: ГННУС (Гигантское Неопознанное Наземное Уникальное Существо)

List of appearances




Video games


  1. 1.0 1.1 Nelson, Arvid (April 24, 2021). Wish I could give you the exact height, but you'd have to check with Legendary – I was just a "hired gun" for them, didn't have access to their data vaults. Same species, yes.. Twitter. Retrieved on April 24, 2021.
  2. 2.0 2.1 MONARCH Declassified: The Godzilla Revelation. Godzilla DVD/Blu-Ray Special Features.
  3. 3.0 3.1 "Monarch Superspecies Profile: MUTO #MonsterverseWatchalong" — @Legendary on Twitter.
  4. 4.0 4.1 "George Tew" - George Tew on Twitter.
  5. 5.0 5.1 New Century Special Effects Movie Guide. Yosensha. pp. 11, 13. 11 January 2015. ISBN: 978-4-8003-0563-3.
  6. Godzilla Movie CLIP MUTO (2014) - David Strathairn, Gareth Edwards Movie HD
  7. Wingard, Adam (April 22, 2021). I'm Adam Wingard, director of Godzilla vs. Kong. AMA!. Reddit. Retrieved on April 22, 2021.
  8. 8.0 8.1 "GODZILLA" by D. Callaham. Revisions by David S. Goyer. Current Revisions by Max Borenstein. 2012-06-20.
  9. Lookdev Queen MUTO - Mike Dougherty Instagram
  10. Concept art for a 4-winged Hokmuto.
  11. Concept art for a 6-appendaged Hokmuto.
  12. intro.mp3
  13. Cox, Greg. Godzilla: The Official Movie Novelization. Titan Books. ASIN: B00MLDU6TO.
  14. https://twitter.com/Legendary/status/1248409968732065792?s=20
  15. https://twitter.com/arvidthetwit/status/1306014254160642051?s=20&t=7QLa3S-jS_K8g5fLcstCcA
  16. https://twitter.com/arvidthetwit/status/1306014621292261376?s=20&t=7QLa3S-jS_K8g5fLcstCcA
  17. https://twitter.com/GodzillaMovie/status/1248410130686734343?s=20
  18. https://rue-morgue.com/sinister-seven-godzillas-mike-dougherty-king-of-the-monsterkids/

Film-based kaiju
Godzilla kaiju
King Kong kaiju
Mothra kaiju
Gamera kaiju
Other kaiju
Scrapped kaiju